Note close correspondence between species cladogram (right) and the species distributions (left). Closely-related Etheostoma veriatum, osburni, and kanawhae occupy the Appalachian highlands, and E. tetrazonum and euzonum the Ozark highlands. Wiley, E.O. and Mayden, R.L. 1985. Species and speiation in phylogenetic systematics, with examples from the North American fish fauna. Annals Missouri Botanical Garden 72: 596-635.